Eyelid
2007 Schools Wikipedia Selection. Related subjects: Health and medicine
An eyelid is a thin fold of skin and muscle that covers and protects an eye. The levator palpebrae superioris muscle retracts the eyelid to "open" the eye. This can be either voluntarily or involuntarily. The human eyelid features a row of eyelashes which serve to heighten the protection of the eye from dust and foreign debris. "Palpebral" means relating to the eyelids.
Anatomy
Layers
In the human eyelid, there are various layers; from superficial to deep, they are: Skin, subcutaneous tissue, orbicularis oculi, orbital septum & tarsal plates, palpebral conjunctiva The meibomium glands lie within the eyelid and secrete tears. They often become blocked leading the blepharitis
Innervation
In humans, the sensory nerve supply to the upper eyelids is from the infratrochlear, supratrochlear, supraorbital and the lacrimal nerves from the ophthalmic branch (V1) of the trigeminal nerve (CN V). The skin of the lower eyelid is supplied by branches of the infratrochlear at the medial angle, the rest is supplied by branches of the infraorbital nerve of the maxillary branch (V2) of the trigeminal nerve.
Blood supply
In humans, the eyelids are supplied by 2 arches on each upper and lower lid. The arches are formed by anastamoses of the lateral palpebral arteries and medial palpebral arteries, branching off from the lacrimal artery and ophthalmic artery, respectively.
Usage
When an eye becomes dry, closing the eyelid and opening it again rapidly — referred to as " blinking" — can help to spread moisture across the surface of the eye and ease the discomfort. Blinking also serves the purpose of helping to remove irritants which have landed in the eye. When a person chooses to blink one eye as a signal to another in a social setting, it is known as " winking."
Most animals with eyelids have a reflex to close the eyes when a threat comes too near. This is done involuntarily to protect the eye from contact with the threat. It is often strong enough to overcome any voluntary resistance.
Eyelids also serve the purpose of helping the animal to control the amount of light entering the eye (control of the iris is autonomic). Without eyelids, many animals would be helpless to block visual sensory overload under extremely bright light. Many animals also use eyelids to block light from reaching the eyes during sleeping cycles.
Types
In humans, each eye has an upper and lower eyelid which operate as a pair, however, it is primarily the upper eyelid that moves across the exposed surface of the eye during blinking. Lower eyelids in most animals move vertically.
Many terrestrial vertebrates have an eyelid known as a nictitating membrane, or haw. This eyelid is closer to the eye than the outer lids and is usually transparent. The purpose of the nictitating membrane is to add protection to the eye from debris and irritation as well as serve as a barrier while swimming to land animals. Camels, Crocodiles, birds, and polar bears are among the others which have this membrane. The nictitating membrane sweeps across the eye diagonally or horizontally. In humans, the plica semilunaris (or semilunar fold) is thought to be the vestigial remnant of the nictitating membrane.
It has been suggested that eyelids evolved as a way to remove debris from the eyes. Given that fish have a constant stream of water flowing over their eyes, it is not surprising that they do not have eyelids or need specialized membranes to perform this function.